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Nothing in Owen’s life has been right since his father died in prison, accused of a crime Owen is certain he didn't commit. Monroe, the IT guy at school, might finally bring Owen the means to clear his father’s name by letting him use an Animus—a device that lets users explore the genetic memories buried within their own DNA. The experience brings Owen more than he bargained for. During a simulation, Owen uncovers the existence of an ancient and powerful relic long considered legend—the Trident of Eden. Now two secret organizations will stop at nothing to take possession of this artifact—the Brotherhood of Assassins and the Templar Order. It soon becomes clear to Owen that the only way to save himself is to find the Trident first.
Under the guidance of Monroe, Owen and a group of other teenagers go into a memory they all share within their DNA: the 1863 Draft Riots in New York City. Owen and his companions will find themselves tested on the gritty streets of New York, and their experiences in the past will have far-reaching consequences in the present.